ISSN 1188-603X

No. 218 March 23, 1999 Victoria, B.C.
Dr. A. Ceska, P.O.Box 8546, Victoria, B.C. Canada V8W 3S2


From: Spribille_Toby/

THE THIRD ANNUAL PACIFIC NORTHWEST BRYOLOGICAL FORAY will be held 13-16 May 1999 at the Cowichan Lake Research Station on south Vancouver Island, British Columbia. This year's bryoforay features an action-packed agenda including an introductory lecture on the basics of bryology for beginners, hands-on laboratory time and field trips to several sites (including Garry oak-madrone woodlands, coastal rainforests and a coastal true bog) on the southern end of Vancouver Island with, Dr. Wilf Schofield, who will accompany us. In addition, we will get the opportunity to share information and recent research through a series of informal lectures and slide shows. If you have anything you wish to present, please contact us to be placed on the agenda.

Cowichan Lake Research Station is located approximately one and a half hours north of Victoria, about five kilometers northwest of the town of Lake Cowichan in south-central Vancouver Island. From the Island Highway between Duncan and Nanaimo, go west on Highway 18 about 29 km to the town of Lake Cowichan, go straight through town, and look for the sign for the Research Station. Turn right on the Forestry Road and follow it out to the end.

The price of the foray includes staying three nights at Cowichan Lake, as well as dinner on the first day, all three meals Friday and Saturday and breakfast and a sack [bag] lunch on Sunday. Transportation to and from the field will not be provided; plan on carpooling or bringing your own transportation. Participants coming from the mainland should be advised of ferry costs from Tsawwassen to Swartz Bay and from Horseshoe Bay to Nanaimo, these typically run $35-$40 (Canadian) one way with one passenger.

Draft agenda (more specific details will be provided upon registration):

Thursday, 13 May
1100-1200 registration
PM Introduction to bryology: lectures and lab session

Friday, 14 May
AM Lecture session
PM Field trip
After dinner in laboratory with samples

Saturday, 15 May
All day field trip
After dinner in laboratory with samples

Sunday, 16 May
AM Field trip
1200 Break up and head home
Price schedules:
Can$120.00 per person for all four days including three nights lodging, three dinners, three breakfasts and three lunches. Registration fees will be collected at the door when the participants arrive. Payments in US Dollars will be assessed based on the exchange rate of that day.
What to bring:
The normal field gear-- boots, raingear, hand lens, etc. Light microscope, slides, cover slips, if you have one. Copy of Lawton's Moss Flora of the PNW, if you have one.
If you would like to present any recent work involving bryophytes as an informal lecture, please let us know the title of your presentation, approximate length, and whether or not you will have slides, overhead projections, etc. We encourage participation in this information sharing event!
Registration will be limited to 30 participants; a waiting list will be maintained to fill any subsequent vacancies on a first-come, first-served basis. To register, please send name, address, phone, fax and e-mail addresses, specify if you have special food needs (e.g., vegetarian), and whether or not you will be bringing a light microscope, to: Judy Harpel,, or phone (360) 891-5121.

REGISTRATION DEADLINE IS 16 APRIL. We look forward to seeing you at Cowichan Lake!


From: Dr. S. Galen Smith []

Many people are confused by changes in the names of the triangular-stemmed ("three-square") bulrushes Scirpus americanus (olneyi) and S. pungens. This confusion will gradually be resolved as more floras (such as the new Jepson Manual - Hickman 1993) using the new names are published. The following explanation may be of some help:

Schuyler (1974) corrected the use of the names for these species as follows:

The type of S. americanus Persoon is conspecific with the type of S. olneyi A. Gray, and S. americanus is the earlier name. Thus the correct name for this species is Scirpus americanus Persoon, synonym S. olneyi A. Gray.

The type of Scirpus pungens Vahl belongs to the species to which Scirpus americanus has long been misapplied. Thus the name Scirpus pungens Vahl is the correct name for this species.

Further, upon the dismemberment of the polyphyletic genus Scirpus, S. americanus becomes Schoenoplectus americanus (Persoon) Volk. ex Schinz & R. Keller, and S. pungens becomes Schoenoplectus pungens (Vahl) Palla.

I am recognizing 3 varieties of Schoenoplectus pungens for the upcoming volume 23 of the Flora of North America (Smith 1995):

Schoenoplectus pungens var. pungens: stigmas 3, achenes lenticular. Eastern North America and Europe. Intergrades extensively with var. longispicatus in the region of sympatry.

Schoenoplectus pungens var. longispicatus (Britton) S.G. Smith. Stigmas 3, achenes often compressed-trigonous. Central and western North America, northern Mexico, temperate South America.

Schoenoplectus pungens var. badius (Presl) S.G. Smith. Like var. longispicatus but floral scales very dark chestnut-colored, and apical notches and awns on floral scales somewhat shorter. Pacific coastal temperate North and South America, Australia, New Zealand, Tasmania.


Hickman, J.C. [ed.] 1993.
The Jepson Manual: Higher plants of California. University of California Press, Berkeley, CA. 1400 p.
Schuyler, A.E. 1974.
Typification and application of the names Scirpus americanus Pers., S. olneyi Gray, and S. pungens Vahl. Rhodora 76: 51-52.
Smith, S.G. 1995.
New combinations in North American Schoenoplectus, Bolboschoenus, Isolepis, and Trichophorum (Cyperaceae). Novon 5: 97-102.


From: Adolf Ceska []

I don't like the idea of switching the name "Scirpus americanus" from the species with blunt stem edges (S. pungens) to that with sharp edges (S. olneyi). In my opinion, the name "Scirpus americanus" should be rejected before we have to return to the pre-Linnean nomenclature in order to understand what we are talking about ("that one with sharp edges" vs. "that one with blunt edges"). If we reject "Scirpus americanus" we will then be dealing with S. olneyi and S. pungens without any confusion and all other combinations based on "Scirpus americanus" will be rejected too. If we treat this group of Scirpus s.lato as Schoenoplectus, we will end up with Schoenoplectus olneyi (A. Gray) Palla and Schoenoplectus pungens (Vahl) Palla.

Article 57.1 of the International Code of Botanical Nomenclature, if I understand it correctly, actually prohibits the use of "Scirpus americanus" for the entity known as S. olneyi unless the proposal of rejecting "Scirpus americanus" is itself rejected:

"57.1. A name that has been widely and persistently used for a taxon or taxa not including its type is not to be used in a sense that conflicts with current usage unless and until a proposal to deal with it under Art 14.1 [i.e., to put it on the Nomina Conservanda list] or 56.1 [i.e., to put it on the Nomina Rejicienda list] has been submitted and rejected."

From this point of view, the use of "Scirpus americanus" for the taxon previously known as S. olneyi (as in Hickman 1993, or in Kartesz 1994) may be against the International Code of Botanical Nomenclature.


Hickman, J.C. [ed.] 1993.
The Jepson Manual: Higher plants of California. University of California Press, Berkeley, CA.
Kartesz, J.T. 1994.
A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. 2nd Edition. Vol. 1+2. Timber Press, Portland, OR.


Geiser, L.H., K.L. Dillman, C.C. Derr, & M.C. Stensvold. 1998.
Lichens and allied fungi of southeast Alaska. Pp. 201-243 in: Glenn, M.G., R.C. Harris, R. Dirig, & M.S. Cole. Lichenographia Thomsoniana: North American lichenology in honor of John W. Thomson. Mycotaxon, Ltd. Ithaca, NY.
You can order this reprint by e-mail:
mstensvold/ (Mary Stensvold) lgeiser/ (Linda Geiser)

A checklist of 508 lichen and allied fungal species with regional habitat, distribution and abundance information has been compiled for southeastern Alaska. The lichen flora of this area is a rich mixture of Pacific Northwest temperate rain forest and Arctic components, and it is enhanced by topographic and habitat variations within the region. Great expanses of old-growth forests and excellent air quality provide habitat for many lichens elsewhere rare or imperiled. 23 species and 1 subspecies are reported new to Alaska and 4 species are new to the United States.

Lichenographia Thomsoniana, published in honor of John W. Thomson, is a collection of 28 important articles devoted to the taxonomy and chorology of lichens in North America. The collection contains, among others, another paper from Alaska (Kodiak Island by S. S. Talbot), a taxonomical paper from the Queen Charlotte Islands (by I.M. Brodo & V. Wirth), and two taxonomical papers co-authored by T. Goward.

The entire volume can be purchased for US$35 from:
Mariette Cole, 3010 West 112th Street, Bloomington, MN 55431


From: Adolf Ceska []

Please, add the following references to "Dr. R.T. Ogilvie - Botanical bibliography" [BEN # 210]:

Ogilvie, R.T. & T. Furman. 1959.
Effect of vegetational cover of fence rows on small mammal populations. Ecology 40: 140-141.
Ogilvie, R.T. & R.C. Scace. 1968.
Guide for field trip to Banff National Park, October 12 and 13, 1968. Pp. 989-1015. Appendix A. in: Nelson, J.G & R.C. Scace. 1968. The Canadian National Parks: Today and tomorrow. Studies in Land Use History and Landscape Change. University of Calgary, Calgary, Alberta. 1027 p.
Ogilvie, R.T. 1969.
Itinerary and guide for field trip in Banff National Park, August 21, 1969. Phytochemical society of North America Annual Meeting, August 1969. 13 p.
Ogilvie, R.T. 1969.
Vegetation zonation in the Rocky Mountains of Alberta, Canada. Abstracts of the papers presented at the XI International Botanical Congress, Seattle, WA, 1969: 162. [abstract]
Geist, V., R.T. Ogilvie, D.H. Gubbe & I.D. Hubbard. 1974.
Report on Wolf Lake, panel 10, c.t. site 18. [University of Calgary, Alberta] ix + 238 p.
Vroom, G.W., S. Herrero & R.T. Ogilvie. 1976.
The ecology of winter den sites of grizzly bears in Banff National Park, Alberta. Pp. 321-330 in Pelton, M.R., J.W. Lentfer, & G.E. Folk [eds.] Bears : their biology and management : a selection of papers from the Third International Conference on Bear Research and Management, held at the 54th Annual Meeting of the American Society of Mammalogists, Binghamton, New York, USA and First International Theriological Congress, Moscow, USSR, June 1974. Morges : International Union for Conservation of Nature and Natural Resources. 467 p.
Scace, R.C. & R.T. Ogilvie. 1978.
Fort Walsh National Historic Park: Period landscape history. Report prepared for Parks Canada, Prairie Region Office, Department of Indian and Northern Affairs. Scace & Associates, Ltd. Calgary, Alberta. 152 p.
Ogilvie, R.T. & A. Ceska. 1988.
Endemic and disjunct vascular flora of the North Pacific Coast. [Abstract] Canad. Bot. Assoc. Annual Meeting, Program & Abstracts, Victoria, B.C.
Ogilvie, R.T. 1992.
Rare and endangered alpine plants in British Columbia. Pp. 131-142 in Rautio, S. [ed.] Community action for endangered species. Report of the Federation of the B.C. Naturalists and Northwest Wildlife Preservation Society, Vancouver, BC. 238 p.
Ogilvie, R.T. 1997.
Vascular plants and phytogeography of Brooks Peninsula. Pp. 5.1-5.48 (Chapter 5) in: Hebda, R. J. & J. C. Haggarty. 1997. Brooks Peninsula: An Ice Age refugium on Vancouver Island. Occasional Paper No. 5, B.C. Ministry of Environment, Lands and Parks, Victoria, B.C.
Hebda, R.J., R.T. Ogilvie, H. Roemer & A. Banner. 1997.
Vegetation of Brooks Peninsula. Pp. 8.1-8.63 (Chapter 8) in: Hebda, R.J. & J.C. Haggarty. 1997. Brooks Peninsula: An Ice Age refugium on Vancouver Island. Occasional Paper No. 5, B.C. Ministry of Environment, Lands and Parks, Victoria, B.C.

The following references were not cited correctly in BEN # 210:
Kirby, C.L. & R.T. Ogilvie. 1969.
The forests of Marmot Creek Watershed Research Basin. Canadian Forestry Service Publication No. 1259, Queen's Printer, Ottawa. 37 p.
Ogilvie, R.T. 1989.
Disjunct vascular flora of northwestern Vancouver Island in relation to Queen Charlotte Islands' endemism and Pacific Coast refugia. Pp. 127-130 in: Scudder, G.G.E. & N. Gessler [eds.] The outer shores. Queen Charlotte Islands Museum Press, Skidegate, B.C. 327 p.

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