FREIRE-FIERRO, ALINA. Department of Biology, University of Missouri-St. Louis, MO 63121-4499. - Revision of Aciotis (Melastomataceae).
Aciotis is a herbaceous genus that includes about 20 species
distributed in Central America, tropical South America and the West
Indies. It occurs in disturbed areas up to 1700 m elevation. Previous
morphological studies of the genus have used anther size, pubescence
type, leaf size, and the width of wings on the stems for species
distinctions. For my revision, I have used additional characters:
habit, number of pairs of paraclades of first order, inflorescence
type, anther color, and presence of fibers in the hypanthium wall.
Fruit characters had not been used before because it was assumed that
Aciotis has only dry capsular fruits. However, field work has
revealed that the genus has representatives with dry capsular fruits
(e.g. A. polystachya, A. annua), species with fleshy capsular
fruits (A. aequatorialis, A. acuminifolia) and species with
berry-like fruits (A. rostellata, A. alata, and A.
purpurascens). The capsular fruits persist on the plant, and the
ovary-hypanthium walls can be caducous or persistent. The berry-like
fruits fall off in their entirety when ripe. Species of Aciotis
can be divided into two groups, which seem to be correlated with
chromosome number and geographic distribution: 1) annuals with lax
inflorescences, capsular and small fruits, low chromosome numbers and
a restricted distribution; and 2) perennials with dense and conical
inflorescences, berry-like and large fruits, higher chromosome numbers
and wide distribution.
Key words: Aciotis, Biogeography, Melastomataceae, Morphology, Neotropics