GOLDMAN, DOUGLAS H.1*, MARK W. CHASE2, AND MICHAEL F. FAY2. 1Department of Botany, University of Texas, Austin, TX 78713; 2Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3DS. - Phylogeny and circumscription in Calopogon (Orchidaceae) as inferred from ITS sequence and AFLP data.
The genus Calopogon consists of five species and three
varieties of terrestrial orchids. Three species, C. barbatus,
C. multiflorus, and C. pallidus are found in coastal
plain savannas of the south-eastern United States. Another species,
C. oklahomensis, recently described, is found in prairies and
savannas from Minnesota south to Texas and Louisiana. The fifth
species, C. tuberosus, consists of three varieties, var.
tuberosus, found in bogs and savannas from Texas and Manitoba,
east to the Atlantic coast, var. latifolius, found in maritime
bogs in eastern Canada, and var. Simpsonii, found in alkaline
savannas in the Bahamas, Cuba, and south Florida. ITS sequences were
obtained for 57 populations within the genus, representing the
geographic and morphological extremes of each taxon. AFLP data was
obtained for 58 plants representing 25 populations within the genus,
one artificial hybrid, and one putative natural hybrid. The 25
populations sampled represent the geographic extremes of all taxa. In
general, there is considerable agreement between the ITS parsimony
analysis and the AFLP UPGMA analysis at the species level, which is
congruent with many significant morphological features. AFLP,
however, shows greater resolution at the population level, although
both data sets generally place sampled populations into previously
defined species. ITS data do not refute the distinctiveness of the
different varieties of C. tuberosus, due to lack of resolution,
whereas AFLP data generally support the varieties. Of particular
interest is the intermediacy that both data sets suggest of C.
oklahomensis. This species is polyploid, while the other four
species are diploid, and numerous morphological features in this
species suggests that it may be of hybrid origin. If available,
comparisons with plastid RFLP data will also be presented.
Key words: AFLP, Calopogon, circumscription, hybrid speciation, ITS, phylogeny