CRAWFORD, PHILLIP AND WAYNE ELISENS.* Department of Botany and Microbiology, University of Oklahoma, Norman, OK 73019. - Genetic variation among native North American toadflaxes (Linaria, Scrophulariaceae).
The Linaria canadensis complex (Linaria section
Leptoplectron or genus Nuttallanthus) comprises a group
of morphologically variable taxa which are recognized at specific or
varietal rank and are native to North America. Linaria
canadensis and L. texana are locally abundant in sandy
soils and appear to be naturalized in other areas of the western
hemisphere. Linaria floridana is narrowly distributed, and is
confined primarily to Atlantic and Gulf coastal areas where all three
taxa occur occasionally in mixed populations. Although species
boundaries are unresolved among New World toadflaxes, previous
morphological studies support hypotheses that the complex forms a
monophyletic group distinct from the 150 Eurasian species of
Linaria. We examined variation within and among 54 populations
of the Linaria canadensis complex at 15 putative isozyme loci
encoding 12 enzyme systems. Chasmogamous flowers have showy blue
corollas and are visited by butterflies and bees, but cleistogamy,
self-pollination, and self-compatibility characterize plants from
investigated populations. High levels of selfing are indicated by
field and greenhouse mating studies and by low levels of
infrapopulation genetic polymorphism and observed heterozygosity.
Analyses of morphological variation differentiate three primary
morphotypes, but isozyme data identify two primary groups of
populations with many qualitative allelic differences and with genetic
identity coefficients less than 0.62: 1) L. canadensis and 2)
L. floridana + L. texana. Isozyme data also indicate
that these two groups are reproductively isolated in areas of sympatry
where mixed populations occur. Population clusters exhibit a
pronounced geographical pattern which may reflect historical
biogeographic isolation and recent migrational pathways.
Key words: Isozymes, Linaria, Scrophulariaceae, species delimitation