LAMMERS, THOMAS G.* AND LINDA GLASS. Department of Botany, Field Museum of Natural History, Roosevelt Road at Lake Shore Drive, Chicago IL 60605-2496. - Morphological variation in the Lobelia polyphylla complex (Campanulaceae) of Chile.
The Lobelia polyphylla complex (Campanulaceae: Lobelioideae)
comprises small (0.6-2 m tall) polycarpic lactiferous shrubs endemic
to the xerophytic dwarf shrub and sclerophyllous shrub regions of
Chile (latitude 27-34S), at elevations from sea-level up to 900
(rarely 1200) m. Flowers are quite uniform within the complex: small,
unilabiate, dark wine-purple in color, and visited by bees, with
anthesis occuring from late winter to early summer (August to
February). In contrast, the sessile deciduous leaves and floral
bracts show an extraordinary range of variation in size and shape,
spanning the entire spectrum from linear laminas 4 mm wide to widely
ovate blades nearly 5 cm across. Based largely on this extreme
variation in foliar features, a total of 13 species and four varieties
referable to the Lobelia polyphylla complex have been
described, but botanical opinion has been divided on how many to
recognize and at what rank. Wimmer's 1953 treatment in Das
Pflanzenreich recognized two species, one of which was subdivided
into four varieties plus four heterotypic forms. However, during
field work in Chile, this classification proved impossible to
implement. To resolve these questions, we performed cluster analyses
and principal components analyses on a data set representing 34
morphological characters extracted from 64 specimens, collected
throughout the geographic and ecological range of the complex. These
analyses supported the recognition of only a single highly polymorphic
but undivided species, Lobelia polyphylla. Further, the
analyses were unable to detect any correlation between morphological
variation and geography, ecology, or elevation.
Key words: Campanulaceae, Lobelia polyphylla, morphological variation, numerical phenetics