ANDERS, CONSTANCE M.*, NICHOLAS DROZDA, AND ZACK E. MURRELL. Department of Biology, Western Kentucky University, Bowling Green, KY 42101. - Genotypic and phenotypic variation in Spiraea virginiana Britton.
Spiraea virginiana is a federally listed rhizomatous shrub
endemic to the southern Blue Ridge and Appalachian Plateau Provinces.
The typical habitat of S. virginiana is along scoured sections
of high gradient streams. No viable seeds or seedlings have been
found in the wild, suggesting that populations within drainages are
products of vegetative reproduction, most probably occurring when
rhizomes broken loose from upstream populations wash downstream to
form new ramets. There is considerable confusion, therefore,
regarding identification of individuals and thus no known mechanism
for evaluating population size and structure. Phenotypic variation in
S. virginiana was investigated through a morphometric
examination of leaf size and shape, using Morphosys to make 25 leaf
measurements. These data were analyzed using Principal Components
Analysis, to identify any morphological variation within and between
drainages. We are currently examining genetic identity across the
distribution using Randomly Amplified Polymorphic DNA (RAPDs), in an
attempt to evaluate the genetic diversity. Genetic uniformity among
the drainages would suggest that either extant populations are the
ancestors of migrants that persisted through the glacial maximum in
the watershed of the Gulf of Mexico, or are the products of a severe
bottleneck during the Hypsithermal Period. Genetic variation among
the drainages would suggest that S. virginiana persisted
through the last glacial maximum within the present day drainage
systems, supporting Ogle's hypothesis that the present distribution of
S. virginiana represents the remnants of a more widespread
distribution in the past. The RAPD data generally show genetic
uniformity within drainages, suggesting that vegetative reproduction
via dislodging of rhizomes during flooding is the predominant means of
dispersal. The morphometric and RAPD data suggest that the northern
part of the range exhibits greater variation than the middle and
southern portion of the range, supporting OgleÕs hypothesis.
Key words: clonal growth, intraspecific genetic variation, morphometrics, Pleistocene migrations, RAPDs, Spiraea virginiana