HERMANN, BRIAN P.*, TARUN K. MAL, RUSSEL J. WILLIAMS, AND NORMAN R. DOLLAHAN. Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, PA 19085-1699 USA. - Quantitative evaluation of stigma polymorphisms in a tristylous weed, Lythrum salicaria(Lythraceae).
Tristyly involves three different forms of flowers that differ
reciprocally in the heights of stigmas and anthers within flowers.
Apart from the style and stamen lengths, heterostylous species also
demonstrate pollen and stigma polymorphisms. We quantified stigma
polymorphisms in tristylous Lythrum salicaria. We measured the
stigma diameters, structure of papillae, density and distribution of
papillae on the stigma from flower samples from 201 individuals
belonging to the three morphs. The diameter of stigma and the
distribution of papillae were quantified under a scanning electron
microscope, and the structure of papillae was determined using a light
microscope. The stigma diameter and the density of papillae in the
long morph were significantly greater than both in the mid and short
morphs, but the total number of papillae per stigma did not differ
across the morphs. The length and diameter of papillae at the apex,
neck and basal regions were significantly greater in the long morph
followed by the mid and short morphs. A discriminant function analysis
separated the long morph from the mid and short morphs based on the
canonical scores of measurements of papillae structure. The
polymorphisms of stigma coupled with those of pollen may play a
functional role in self-incompatibility mechanisms.
Key words: Heterostyly, incompatibility, Lythrum salicaria, purple loosestrife, scanning electron microscopy, stigmatic papillae.