Multiple molecular data based phylogenies support a relationship among Platanaceae, Nelumbonaceae, and Proteaceae. Developmental and anatomical results are interpreted in a metameric framework to clarify the organizational architecture of Platanus. These studies clarify the organization of different leaves/pherophylls (hülle), ocreas and the development of axillary meristems in intrapetiolar positions. Based on metameric concepts, the platanoid architecture is composed of a conserved repeating growth unit (RGU) composed of six - eight metamers. Each RGU generally terminates with the production of an inflorescence. There is variation among the metamers of an RGU depending on their position along the axis. Integrated into a phylogenetic comparative framework, these data provide a better understanding regarding the architectural transformations associated with Platanus and its putative relatives.

Key words: hulle, ocreas, phylogenetics, Platanaceae, vegetative architecture