LI, XIAOJIE*, J.M. BASKIN, AND C.C. BASKIN. School of Biological Sciences, University of Kentucky, Lexington, KY 40506. - Morphology and physiology of fruit and seed growth and development in two bird-dispersed shrubs, Rhus aromatica, and R. glabra (Anacardiaceae).
Morphology and physiology of fruit and seed development were compared
in wild populations of two shrubs, Rhus aromatica and R.
glabra (Anacardiaceae) in 1996 and 1997. Flower buds of R.
aromatica formed in August, and plants flowered and set fruits the
following April and June, respectively, whereas R. glabra
formed flower buds after the plants leafed-out in early June and
flowered in late-June to early-July, one week after the endocarp of
R. aromatica fruits had become impermeable to water. However,
the two species showed strong similarities in fruit and seed
development. It took only about 2 months for their flowers to develop
into drupes. The single sigmoidal growth curve in for inrease in fruit
size and dry weight of both species differs from the cyclic
(double-sigmoidal) one described for typical drupes such as peach and
cherry. The order of attainment of maximum size was fruit and endocarp
(same time), seedcoat, and embryo, each separated by 1 week. By the
time fruits turned ripe-red, the embryo had reached full size and
become germinable. The endocarp was the last fruit component to reach
physiological maturity, which coincided with the development of
impermeability and a moisture content of < 15%. At this time, about
50%, 37%, and 13% of the dry weight of the drupe was allocated to the
exocarp plus mesocarp unit, endocarp, and seed, respectively. This
study lends support to the opinion of some plant taxonomists that
Lobadium, of which R.aromatica is the type species, should be
considered a subgenus of Rhus, rather than a distinct genus, as
advocated as early as Rafinesque's time and defended by Barkley in
1965.
Key words: fruit, morphology, physiology, Rhus aromatica, Rhus glabra, seed