DUDLE, DANA A. Department of Biology, Indiana University, Bloomington, IN 47405. - Ecological and morphological factors contributing to maintenance of male-sterility in gynodioecious populations of Lobelia siphilitica (Lobeliaceae).
In theory, male-sterility may be maintained in gynodioecious
populations through genetic factors only. However, it is possible
that ecological factors, such as different habitat requirements of the
two sexual morphs, could contribute to or detract from the ability of
male-sterile mutations to invade and spread in populations. Here I
present results showing morph-specific pollinator limitation in a
natural population of Lobelia siphilitica, such that females
are more likely to set fruit than hermaphrodites when pollinator
service is low. The difference in fruit set is apparently caused by
differences in floral morphology: because their stamen tubes contain
no pollen, female flowers are far more likely than hermaphroditic
flowers to become receptive to pollination without several prior
visits from pollinators serving to remove pollen. In addition,
hermaphroditic progeny of females, that definitely carry the
male-sterility factor, enter the pistillate phase more easily than
progeny of hermaphrodites that may not carry the male-sterility gene.
This difference may be a result of hermaphrodites from female lines
containing less pollen than those from hermaphroditic lines, possibly
causing less blockage of the style as it grows through the stamen
tube. Thus, interactions between floral morphology and an ecological
factor, limited pollinator service, may allow the maintenance or
spread of male sterility in some populations of Lobelia
siphilitica.
Key words: dichogamy, floral longevity, gynodioecy, Lobelia siphilitica (Lobeliaceae), male-sterility, pollinator limitation