WONG, K.C.* AND M. SUN. Population Genetics Laboratory, Department of Zoology, University of Hong Kong, Hong Kong.. - Population genetic structure and reproductive biology of Goodyera procera (Orchidaceae): conservation implications.
Goodyera procera (Kew-Gawl.) Hook is an endangered terrestrial
orchid in Hong Kong. Pollination experiments showed that the species
is a self-compatible outbreeder, but dependent on pollinators for
fruit setting. Bagged plants produced no fruits, and artificial
pollinations resulted in 92% fruit set through selfing, 94% through
geitonogamous pollination, and 95% through xenogamous pollination;
whereas fruit set in the open-pollinated control was 75% at the same
sites. Allozyme electrophoresis and random amplified polymorphic DNA
(RAPD) were used to evaluate genetic structure of 15 natural
populations. Despite its outbreeding system, allozyme data revealed a
low level of genetic variation within populations (P = 21.78%, A =
1.22, and H = 0.072) compared with other animal-pollinated outbreeding
plant species. At the population level, RAPD variation was P = 55.13%
and H = 0.18. At the species level, allozyme variation was low with P
= 33%, A = 1.33, and HT = 0.15, and from RAPD data P =
97.03% and HT = 0.29. However, genetic differentiation
among populations was much above average for outbreeding species
(GST = 0.52 and I = 0.909±0.049 based on allozyme data, and
GST = 0.39 and I = 0.859±0.038 based on RAPD data),
suggesting that gene flow was limited among sites. Based on this
information, the following questions were addressed: 1) How did the
species deviate from panmixia in the natural environments? 2) What
were the likely causes (genetic drift, geographical isolation,
anthropogenic disturbance) for the unexpectedly low level of allozyme
variation? and 3) What strategies should be adopted for the protection
and management of this species in Hong Kong?
Key words: allozymes, conservation genetics, Goodyera, RAPD, wild orchids