WHITE1, GEMMA M*, WAYNE POWELL1, AND DAVID BOSHIER2. 1 Cell and Molecular Genetics Department, Scottish Crop Research Institute, Invergowrie, Dundee, DD2 5DA, UK. 2 Oxford Forestry Institute, Oxford University, Oxford, UK. - The dynamics of pollen flow detected in a fragmented population of Swietenia humilis (Zucc.) using SSRs as a marker system.
The effects of tropical deforestation and forest fragmentation on the
population dynamics and the consequent genetic conservation value of
these remnant tree stands is a subject of debate. This work describes
the effect of fragmentation on the levels of diversity and the
patterns of gene flow between 4 remnant stands of Swietenia humilis
(Zucc.): one of the 3 economically important species commonly
known as mahogany. Native to the Pacific coast of Central America, the
once continuous dry forest is now highly disturbed and fragmented by
human and agricultural encroachment. The levels of diversity and the
dynamics of pollen flow were determined using simple sequence repeats
(SSRs) as a marker system. These highly variable markers detected very
high levels of allelic polymorphism with a mean heterozygosity of 0.52
(range 0.04-0.87). The historical population structure
pre-fragmentation, assessed by an analysis of molecular variance
(AMOVA), showed that 98.3% of variation is found within fragments
indicating low population differentiation (Rst=0.02).
Bulked progeny arrays of selected trees from each fragment were
genotyped at 4 SSR loci and a paternity exclusion analysis was
performed on the resulting multi-locus combinations of alleles. It is
evident that the immediate effects of fragmentation on the patterns of
pollen flow is dependent on fragment size and structure. The largest
fragment in this study (n=98) is a good example of 'continuous' forest
and the patterns of pollen flow show a favouring for near neighbour
interactions. But the other smaller fragments of lower tree density
show that there is extensive pollen flow between stands with
distances of up to 6 km. Thus indicating that habitat fragmentation
of S. humilis over this scale does not cause the reproductive
isolation of the trees and that extensive pollen flow provides
connectivity between the patches of remnant forests.
Key words: paternity analysis, pollen flow, simple sequence repeats, Swietenia humilis , tropical dry forest fragmentation