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Courses in Biology

The department offers courses which are slashlisted so undergraduate students may take an undergraduate 4000-level course while graduate students may take a graduate 5000-level course. The lectures in a slashlisted course are the same. However, students in the 5000-level course have substantial additional requirements beyond those for students in the 4000-level course. These additional requirements are listed in the slashlisted course syllabus.

Many of  these courses are not regularly scheduled. If you are interested in a biology course that is not listed below or have questions please contact the Biology Department via email: biology@ou.edu

1003 Contemporary Issues in Biology. An introduction to biology, focusing on the scientific background needed to understand selected issues related to cells, genetics and inheritance, evolution and ecology. Not open to students with credit for 1005 or 1134, or Botany 1114, Zoology 1114 or Zoology 1124. Cannot be used for major credit in Zoology, Botany, or Microbiology. (F, Sp) [II-NL]

1005 Concepts in Biology. Prerequisite: None, but high school or college chemistry is recommended. An introduction to the life sciences, focusing on the structure and function of organisms and their relationship to the environment. Fulfills general education laboratory science requirement. Not open to students with credit for 1003 or 1134, or Botany 1114, Zoology 1114 or Zoology 1124. Cannot be used for major credit in Zoology, Botany, or Microbiology. Laboratory (F, Sp) [II-LAB]

1013 Introduction to Biology. Introductory survey of the fundamental concepts that underlie biological phenomena from the cellular to the ecosystem level. Not open to students with credit for 1003, 1005, or 1134, or Zoology 1114, or Zoology 1124. Cannot be used for major credit in Zoology, Botany or Microbiology. (F, Sp) [II-NL]

1114 Introductory Zoology. Major biological principles and concepts as illustrated in the structure, function and evolution of animals. Emphasis is on self-regulatory mechanisms, especially in the vertebrates, and their adaptive significance. (F, Sp, Su) [II-NL] [HONORS ONLY]

1121 Introductory Zoology Laboratory. Prerequisite: previous completion or concurrent enrollment in 1114. Laboratory study of structure and development of organ systems. Experiments on physiological process of selected vertebrates and invertebrates. (F, Sp, Su) [II-LAB] [HONORS ONLY]

1124 Introductory Biology: Molecules, Cells, and Physiology. Content is focused toward life science majors. Major principles and concepts are presented in the function and physiology of animals, plants, fungi and microbes. Emphasis is on biological chemistry, cell structure and function, cellular energetics, molecular genetics, homeostasis and physiology. Includes biological laboratory experience with emphasis on critical thinking and problem solving, and topics include biochemistry, molecular genetics, cell processes and physiology. Laboratory (F) [II-LAB]

1134 Introductory Biology: Evolution, Ecology and Diversity. Prerequisite: Life science majors only. Major biological principles and concepts as illustrated in a survey of the diversity, behavior, and ecological functions of animals, plants, fungi, and microbes. Emphasis is on evolution, ecology, and diversity. Will include biological laboratory experience with emphasis on problem solving. Problems will be derived from topics in evolution, ecology, and diversity. Will include training in scientific procedures, including laboratory technical skills, writing skills, and introduction to statistical analysis. Recitation will include discussion and case study analysis of the major biological principles presented in the lecture within the context of health and the environment. Will involve problem sets, primary journal articles, and writing assignments. Includes both online and classroom activities. Field trips. Laboratory (F, Sp)

1203 The Age of Dinosaurs (Crosslisted with Geology 1203). Introduction to basic principles and theories in zoology (evolution, systematics, vertebrate morphology and relationships) and geology (geologic time, earth history, plate tectonics, sedimentation and stratigraphy), focusing on the evolutionary history of Dinosauria. May not be counted for major coursework in either department or for general education requirements. (Irreg.) [II-NL]

2011 Critical Evaluation of Biological Research. Prerequisite: 1114 and 1121, or 1124, and 2012 or concurrent enrollment. Emphasis will be placed on understanding the scientific method, how it applies to biological research, and implications for biomedical issues. The course will also introduce students to research in the four core areas of study in the Zoology department (physiology, ecology and systematics, animal behavior, and cellular, genetic, and developmental biology) while emphasizing three basic themes in biology (homeostasis, biological diversity, and evolution). (Irreg.)

2012 Foundations in the Biomedical Sciences. Prerequisite: 1114 and 1121, or 1124. An introduction to professional aspects of the biomedical sciences including basic biostatistics and data interpretation, critical evaluation of scientific literature, and discussion of case studies in biomedical ethics. Emphasis will be placed on the medical relevance of important generalizing themes in biology, including evolution, homeostasis, and biodiversity. Critical thinking and evaluation of current issues in medicine and medical research will be incorporated into class activities. (Irreg.)

2013 Evolution. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and Biology 1134. Process of evolutionary change. Topics include origin of species, evolution above the species level, major transitions of life on earth, application of evolutionary principles in medicine and conservation. (F, Sp)

2103 Everyday Evolution. Prerequisite: sophomore standing. Overview of processes of biological evolution, with emphasis on how evolution affects daily life (e.g., in agriculture, medicine, the law and social interactions). Process of speciation and extinction examined within the context of life’s past history on earth and prospects for its future. Not available to Botany, Microbiology or Zoology major credit. (Irreg.) [II-NL]

2124 Human Physiology. Prerequisite: 1114 and 1121 with a grade of C or better, or 1124 with a grade of C or better; a course in chemistry or physics. May not be applied for zoology major credit. Open only to majors in nursing, physical therapy, health and exercise science, and selected fields. Function of vertebrate organ systems in homeostasis. Circulation, digestion, endocrine and nervous control, metabolism, muscle action and respiration, with emphasis on humans. Laboratory (F, Sp, Su)

2234 Introduction to Human Anatomy. Prerequisite: 1114 and 1121 with a grade of C or better, or 1124 with a grade of C or better, sophomore standing, and permission of department. (Cannot be enrolled concurrently with 2255). An introduction to the gross morphology of the human body. The course will use a lab/lecture format with extensive use of models, videos, and computer-assisted instruction as well as prosected cadavers. Not for zoology major credit. (F)

2255 Human Anatomy. Prerequisite: 1114 and 1121 with a grade of C or better, or 1124 with a grade of C or better, sophomore standing, and permission of department. (Cannot be enrolled concurrently with 2234.). Open only to majors in health and exercise science, physical therapy, nursing and selected fields. The development and gross morphology of the human body and its systems. Laboratory dissection of human cadavers. Not for zoology major credit. Laboratory (F, Sp)

2404 Ecology and Environmental Quality (Crosslisted with Botany 2404). Prerequisite: sophomore standing. Study of ecological principles and their applications to human systems, study of population, air pollution, water pollution, energy issues, etc. Laboratory exercises focus on learning scientific methods of measurement of environmental quality factors. Laboratory (Irreg.) [II-LAB]

2913 Introduction to Quantitative Biology. Prerequisite: 1114 and 1121, or 1124, or Biology 1134, Mathematics 1523 or 1643 or higher, or permission of instructor. The connections between basic mathematics and how biological data are organized, tested, and interpreted. Includes review of probability theory, introduction to parametric and non-parametric biostatistics, fundamentals of experimental design, and sketches of how optimality theory can be used to generate biological questions. (Sp even-numbered years)

3043 Sensory Functions (Crosslisted with Psychology 3043). Prerequisite: 1114 and 1121, or 1124, or Psychology 1113. A comparative survey of the anatomy and physiology of visual, auditory and olfactory systems, as related to their biological significance. Topics covered include sensory functions in communication, predator avoidance and prey capture. (Irreg.)

3054 Invertebrate Zoology. Prerequisite: ZOO 1114 and ZOO 1121, or BIOL 1134. A survey of the invertebrate animals featuring their classification, morphology, life history, ecology, conservation considerations and evolution. Laboratory (Irreg.)

3063 Veterinary Entomology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. The study of insects and their near relatives, such as ticks, as they relate to the causation of economic loss and transmission of disease organisms in livestock and companion animals. Insect biology, disease transmission, and methods of control will be stressed in lecture. Laboratory emphasizes 1) collection, preservation, and pathogens, and 2) toxicological methods used to control and evaluate insecticides and acaricides. Laboratory (December Intersession)

3073 Medical Entomology. Prerequisite: 1114 and 1121, or Biology 1134. Medical entomology investigates the relationship of insects and other arthropods to the health of humans, domestic animals, and wildlife. Laboratory. (Sp)

3083 Animal Behavior (Crosslisted with Psychology 3083). Prerequisite: 2013 or permission of instructor. Animal behavior from an evolutionary perspective. The effects of natural selections on mechanisms underlying behavior and on diversity of behavior among and within species. (F, Sp)

3092 Animal Behavior Laboratory (Crosslisted with Psychology 3092). Prerequisite: junior standing; concurrent or previous enrollment in 3083. Laboratory and field studies on the adaptive nature of animal behavior, illustrating basic principles of experimental design, data analysis, and scientific writing.Field trips. Laboratory (Irreg.)

3101 Principles of Physiology Lab. Prerequisite: 3103 or concurrent enrollment. Provides students with an introduction to the methods and procedures used in physiological research. Topics covered include data acquisition, analysis and basic statistics, effects of temperature on living systems, nervous system functions, muscle mechanics and physiology, and studies of metabolic rates. In addition to hands-on laboratory experience, library projects and written research papers are used to introduce students to methods of scientific communication. Laboratory (F, Sp)

3103 Principles of Physiology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134, or permission of instructor. One semester of physics and organic chemistry strongly recommended. Introduction to basic concepts of physiology; relation of functions of organisms to physical and chemical principles, and to the environment; discussion of experimental design, constituents of tissues, energy, growth, homeostasis, cellular and organ functions. (F, Sp)

3113 Cell Biology (Crosslisted with Botany, Microbiology 3113). Prerequisite: 1114, or 1124, or Biology 1134, or Botany 1114, and Chemistry 3053. Introduction to the cell as a unit of life. A chemical and physical comparison of procaryotic and eucaryotic cells to include a discussion of cell metabolism, types of metabolic regulation, and an analysis of ultrastructure. Emphasis will be placed on the dynamic changes in metabolism and ultrastructure which occur during the life of a cell. (F, Sp)

3201 Animal Development Lab. Prerequisite: 3203 or concurrent enrollment. Laboratory study of the development and embryology of a variety of animals. Developmental concepts and mechanisms will be illustrated through the use of prepared materials and hands-on experiments. Laboratory (F, Sp)

3203 Animal Development. Prerequisite: ZOO 1124, or ZO 1114 and ZOO 1121; and ZOO 3333; and BIOL 1134. Study of animal development from gamete formation through organogenesis and postembryonic phases in different animal models. Concepts and mechanisms at the tissue, cellular and molecular levels will supplement descriptive analyses of development. (Sp)

3214 Comparative Vertebrate Anatomy. Prerequisite: 1114 and 1121, or equivalent. A study of the anatomy and evolutionary development of vertebrate organ systems. Representative vertebrates are studied in laboratory. Laboratory (Irreg.)

3333 Genetics (Crosslisted with Botany 3333). Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; Biology 1134 recommended. Principles of inheritance at gene, chromosome, and population levels; nature of the genetic material and its involvement in the determination of structure and function. (F, Sp)

3342 Genetics Laboratory (Crosslisted with Botany 3342). Prerequisite: 3333 or concurrent enrollment, or equivalent. The demonstrations, crosses and experiments are designed to illustrate various genetic phenomena, including Mendelian laws, recombination, mutation, natural and artificial selection, and interaction of genotype with environment. The primary organism studied is Drosophila, with some use of corn, Neurospora, and others. Laboratory (F)

3403 Principles of Ecology. Prerequisite: 1114 and 1121, or 1134, or PBIO 1114. Patterns of environments and biological communities; the processes maintaining these patterns. Laboratory (F, Sp)

3440 Mentored Research Experience. 0 to 3 hours. Prerequisites: ENGL 1113 or equivalent, and permission of instructor. May be repeated; maximum credit 12 hours. For the inquisitive student to apply the scholarly processes of the discipline to a research or creative project under the mentorship of a faculty member. Student and instructor should complete an Undergraduate Research & Creative Projects (URCP) Mentoring Agreement and file it with the URCP office. Not for honors credit. (F, Sp, Su)

3960 Honors Reading. 1 to 3 hours. Prerequisite: admission to Honors Program. May be repeated; maximum credit six hours. Will consist of topics designated by the instructor. The content will emphasize work not presented in other courses. (F, Sp, Su)

3970 Honors Seminar. 1 to 3 hours. Prerequisite: admission to Honors Program. May be repeated; maximum credit six hours. Discussion of recent and current research trends and significant developments in zoology. (Irreg.)

3980 Honors Research. 1 to 3 hours. Prerequisite: admission to Honors Program. May be repeated; maximum credit six hours. Will provide an opportunity for the gifted honors candidate to work at a special project under the guidance of a professor in the student's field. Laboratory (F, Sp, Su)

3990 Independent Study. 1 to 3 hours. Prerequisite: one course in general area to be studied; permission of instructor and department. May be repeated; maximum credit six hours. Contracted independent study for topic not currently offered in regularly scheduled courses. Independent study may include library and/or laboratory research and field projects. (F, Sp, Su)

4034 Mammalogy. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. Classification, distribution and natural history of mammals with emphasis on Oklahoma species. Mammals are collected and prepared for scientific collections. Field trips. Some overnight camping. Laboratory. (F even-numbered years)

4044 Ornithology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; andBIOL 1134. Biology of birds. Identification of birds in North America with emphasis on Oklahoma; relationships, natural history and behavior or birds. Field trips. Laboratory (Sp odd-numbered years)

4053 Forensic Entomology. Prerequisite:ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. Lecture will explore the use of insects in the science of forensic entomology and its impact on death scene investigation, neglect or abuse; contamination of food products and other marketable goods and subsequent litigation. Lab will be centered on a "death scene investigation" in which students will collect data from a pig carcass to determine factors that affect the rate of decomposition. Laboratory (May Intersession)

4073 General Entomology. Prerequisite: Sophomore standing, and ZOO 1114 and ZOO 1121, or BIOL 1134, or BOT 1114, or BIOL 1005, or permission of instructor. Introduction to the world of insects. Morphological and physiological adaptations; taxonomy, life histories, and methods of collection. Field trips. Laboratory (F)

4083 Herpetology. Prerequisite: 2013 and Biology 1134 or permission of instructor. An introduction to the study of amphibians and reptiles. Taxonomy, ecology, behavior and life histories of amphibians and reptiles, with emphasis on local forms. Field trips. Laboratory (F odd-numbered years)

4093 Behavioral Ecology. Prerequisite: 3083 or permission of instructor. Ecological basis of animal behavior and the role of behavior in adaptation, speciation, and co-evolution. Topics include behavioral genetics, decision making, sexual selection and sexual conflict, sociality, and human behavior and ecology. (Irreg.)

4113 Cellular Pathology (Slashlisted with 5113). Prerequisite: 3113 or permission of instructor. The course focus is on the molecular and cellular bases of disease and alterations in cellular processes that lead to the development of various pathological conditions. Topics include symptoms of cellular disease, pathology of organelles, cell injury, cell death, immunopathology, neoplasia and genetic disorders. No student may earn credit for both 4113 and 5113. (F)

4153 Endocrinology (Slashlisted with 5153). Prerequisite: 3103; 3113 strongly recommended. The major emphasis of the course is how hormones work at the molecular, cellular, and physiological levels. The course examines the molecular biology of hormonal mechanisms and the integration of those mechanisms to maintain crucial homeostatic control systems. Hormones and their receptors are considered in terms of chemistry, biosynthesis, and mechanisms of action. The course focuses primarily on vertebrate hormones. No student may earn credit for both 4153 and 5153. (Irreg.)

4172 Cellular and Molecular Techniques (Slashlisted with 5172). Prerequisite: ZOO 3113 or permission of instructor. A hands-on introduction to molecular techniques used in modern cell/molecular laboratories and discussed in cell biology courses. No student may earn credit for both ZOO 4172 and ZOO 5172. (Irreg.)

4193 Life History (Slashlisted with 5193). Prerequisite: senior standing or permission of instructor. Overview of theory and empirical studies of life history evolution. Includes lecture, discussion of the primary literature, and analysis of some basic life history problems. No student may earn credit for both ZOO 4193 and 5193 (Irreg.)

4204 Vertebrate Paleobiology (Slashlisted with 5204; Crosslisted with Geology 4204). Prerequisite: 1114 and 1121, or 1124 or Biology 1134; 3214; or permission of instructor. Field trips. Systematics, relationships, zoogeography and evolutionary morphology of the major groups of vertebrates. No student may receive credit for both 4204 and 5204. (Irreg.)

4213 Neurobehavioral Development. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. Specific developmental issues will be discussed from both the behavioral and biological levels of analysis. Cannot be taken for credit if 6213 has been previously taken. (Irreg.)

4223 Cellular and Molecular Neurobiology (Slashlisted with 5223). Prerequisites: 3103, 3113, 3203, or 3333, or permission of instructor. Introduction to cellular and molecular neurobiology through reading and discussion of landmark research papers in five current areas of research. Emphasis will be on understanding modern cellular and molecular research methods and on critical interpretation of scientific data. No student may earn credit for both 4223 and 5223. (Irreg.)

4233 Neurobiology of Disease (Slashlisted with 5233). Prerequisite: ZOO 4833 (preferred), or ZOO 3113 and ZOO 3333, or ZOO 2124 or ZOO 3103; or permission of the instructor. Cellular and molecular mechanisms underlying both normal neuronal function and neuronal disorders. Includes a review of basic concepts in neuroscience through traditional lectures, and reading and discussion of original research articles. Students are required to give oral presentations, write critiques and term papers, and present research posters. No student may earn credit for both ZOO 4233 and ZOO 5233. (F)

4244 Animal Histology. Prerequisite: ZOO 3103 and ZOO 3113, or permission of instructor. Structure and function of animal tissues with emphasis on the cellular basis of tissue and organ function. Laboratory emphasizes the identification of cells and tissues with the use of the light microscope. Laboratory (Sp)

4253 Evolution of Development (Slashlisted with 5253). Prerequisite: 3013 and 3203, or permission of the instructor. Introduction to evolutionary developmental biology (EVO DEVO), study of the developmental basis of evolutionary changes and how developmental mechanisms and developmentally important genes evolve. Latest techniques and methods in evolutionary biology, embryology, developmental biology, molecular genetics, population biology, phylogenetics and genomics. No student may earn credit for both 4253 and 5253. (Irreg.)

4353 Molecular Techniques for Field Biology (Slashlisted with 5353). Prerequisite: 1114 and 1121, or 1124 and permission of instructor; 3333 or 3403 recommended. Selected protocols and data interpretation using molecular techniques to study protein and DNA variation in natural populations and the application of molecular techniques to research problems in ecology, systematics, animal behavior, conservation biology, and related areas. Graduate students enrolled in 5353 will have additional project expectations and written work. Taught at the OU Biological Station. No student may earn credit for both 4353 and 5353. Laboratory (Su)

4383 Functional Genomics (Slashlisted with 5383). Prerequisite: 3113 and either 3203 or 3333 or 4843 or Chemistry 3653 or permission of the instructor. Introduction to how to study the functions (broadly including biochemical activity, cell biological function or organismal function) of all the genes in a genome. Latest techniques and methods in genetics, molecular biology, microarrays, proteomics and computation biology. No student may earn credit for both 4383 and 5383 (Irreg.)

4423 Stream Ecology (Slashlisted with 5423). Prerequisite:ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134; junior or senior standing, or permission of instructor. A combined lecture/laboratory course that focuses on the physical, chemical, and biological features of stream ecosystems, including current theories explaining species interactions and stream function. Course requirements/evaluation including a midterm and final examination, individual research papers and presentations, participation in group laboratory and field experiments, and reading and discussing the primary literature. No student may earn credit for both 4423 and 5423. Laboratory (F-odd numbered years)

4463 Ecology of Lakes. Prerequisite: 3403 or Botany 3453, Chemistry 1315, Chemistry 1415, Physics 1114, Physics 2414 or Physics 2514, or permission of instructor. An introduction to the biology, chemistry, physics and geology of freshwater environments, with emphasis on ecology. (Spring even-numbered years)

4471 Ecology of Lakes Laboratory. Prerequisite: 4463 or concurrent enrollment. Experience in the use of the basic limnological methods and application of these methods to a variety of freshwater environments. Field trips. Laboratory. (Spring even-numbered years)

4493 Ichthyology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. Taxonomy, morphology, ecology and distribution of fishes, with emphasis on those of the region. Laboratory. (F-even numbered years)

4563 Biological Conservation (Slashlisted with 5563). Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and ZOO 2013 or ZOO 3083 or ZOO 3403; and BIOL 1134. Using primary literature, discussion of ecological, legal, and societal issues affecting biodiversity in North America and globally. Topics range from organism centered approaches to conservation to broader implications of alterations of global processes such as elemental cycles and atmospheric temperatures. No student may earn credit for both 4563 and 5563. (F-even numbered years)

4573 Conservation Genetics (Slashlisted with 5573). Prerequisite: 3333 or permission of the instructor. This lecture/discussion course will examine the use of population genetic/ecological genetic principles in the study and management of populations of threatened and/or endangered species. No student may earn credit for both 4573 and 5573. (Sp)

4653 Parasitology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134. Parasitology is an introduction to the biological relationships known as parasitism.  Although there are many different types of parasites, this course will focus on primarily protozoan and helminthes parasites that cause disease in humans and domestic animals. Laboratory (Sp odd-numbered years)

4663 Advanced Limnology (Slashlisted with 5663). Prerequisite: 4423 or 5423, or 4463 and 4471, or permission of instructor. May be repeated with change of content; maximum credit 6 hrs. detailed study of fundamental or contemporary topics within limnology, such as biogeochemistry, nutrient cycling, ecological stoichiometry, biodiversity, and predator-prey and food-web dynamics in aquatic communities. No student may earn credit for both 4663 and 5663 on the same topic. (Sp)

4753 Principles of Phylogenetics (Slashlisted with 5753). Prerequisite: ZOO 2013 and ZOO 3333. Theory and practice of inferring evolutionary history from molecular and morphological data. Applications of the phylogenetic approach in systematics, comparative biology, molecular evolution, and genomics will be covered. No student may earn credit for both 4753 and 5753. (Irreg.)

4813 Hormones and Behavior (Slashlisted with 5813). Prerequisite: 3083 or 3103 or permission of instructor. The relationship between hormones and behavior from an evolutionary perspective. Focus on reproductive and social behavior, with coverage of biological rhythms, food and water intake, and learning. No student may earn credit for both 4813 and 5813. (Irreg.)

4833 Introduction to Neurobiology (Slashlisted with 5833). Prerequisite: 3103 or 3113 or permission of instructor. Introduction to cellular and behavioral neurobiology. Topics covered will include cellular neurobiology, neurophysiology, neuroanatomy, sensory processing, movement, and neurobiology of behavior. No student may earn credit for both 4833 and 5833. (F)

4843 Introduction to Molecular Biology (Crosslisted with Botany and Microbiology; Slashlisted with 5843). Prerequisite: 1114 or 1124, or Botany 1114, or Microbiology 3813 and 3812, and one course in organic chemistry. Introduction to the characteristics and biological functions of nucleic acids and proteins in living cells with emphasis on nucleic acid replication, transcription, translation and regulation; also emphasis on the molecular aspects of microbial genetics—transformation, transduction and conjugation; and emphasis on molecular immunology and genetic engineering/recombinant DNA technology. No student may earn credit for both 4843 and 5843. (F)

4853 Neurobiology of Memory (Crosslisted with Psychology 4853). Prerequisite: 3103 or Psychology 3073. Advanced seminar dealing with current research on how the nervous system responds to, and is changed by, experience. Experience is broadly defined and may include learning, maturation and/or pathology.

4863 Neural Control of Movement (Slashlisted with 5863). Prerequisite: ZOO 3103 or ZOO 3113, or ZOO 4833/ZOO 5833, or permission of instructor. Introduction to neural control of movement through reading and discussion of key original research articles from the 19th century to the present. Students lead discussions and write essays addressing a general question, utilizing data from the articles; students in 5863 write an additional essay. Topics include localization of function, sensory vs. central contributions, roles of single neurons, effects of neuromodulators, and motor learning. No student may earn credit for both 4863 and 5863. (Sp)

4871 Current Topics in Neurobiology (Slashlisted with 5871). Prerequisite:  Permission of the instructor.  May be repeated; Maximum credit three hours.  A seminar course designed to develop a student’s abilities to interpret and critically evaluate research in cellular and behavioral neurobiology.  Involves both public seminars and journal club style discussions of contemporary literature.  No student may earn credit for both 4871 and 5871.  (F, Sp)

4893 Behavioral Neurobiology (Slashlisted with 5893). Prerequisite: ZOO 3103 or ZOO 3113, or ZOO 4833/ZOO 5833, or permission of instructor. Examines neurobiological mechanisms of natural animal behaviors (i.e. neuroethology), utilizing textbook and lectures as well as in-depth reading, discussion, and student presentation of original research articles. No student may earn credit for both 4893 and 5893. (Irreg.)

4913 Quantitative Biology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134; a course in statistics recommended. Techniques for complex data analysis and experimental design. No laboratory. (F)

4960 Directed Readings. 1 to 4 hours. Prerequisite: good standing in University; permission of instructor and dean. May be repeated; maximum credit four hours. Designed for upper-division students who need opportunity to study a specific problem in greater depth than formal course content permits. (Irreg.)

4961 Undergraduate Seminar. Prerequisite: Senior standing in Zoology or permission of department. May be repeated; maximum credit two hours. Survey of current research programs in environmental biology, cell biology, physiology, animal behavior and other fields presented in weekly public seminars by visiting and local experts in biology. (F, Sp)

4970 Special Topics in Zoology. 1 to 3 hours. Prerequisite: three courses in general area to be studied; permission of instructor, department. May be repeated with change of topic; maximum credit nine hours. Seminar or special topic course; may include laboratory or field work. (F, Sp, Su)

4983 Senior Seminar. Prerequisite: ZOO major with senior standing, or permission. An interdisciplinary approach will be used to synthesize ideas from the major fields of zoology. Readings and discussion will focus on contemporary social, ethical and economic issues. (F, Sp) [V]

4990 Independent Study. 1 to 3 hours. Prerequisite: three courses in general area to be studied; permission of instructor and department. May be repeated; maximum credit six hours. Contracted independent study for topic not currently offered in regularly scheduled courses. Independent study may include library and/or laboratory research and field projects. (F, Sp, Su)

5093 Evolutionary Ecology. Prerequisite: graduate standing, or ZOO 2013 or permission of instructor. Study of ecological forces important in evolutionary change in plants and animals. Evolution of sex, breeding systems, life history, speciation, coevolution of animal/plant interactions. (Irreg.)

5113 Cellular Pathology (Slashlisted with 4113).  Prerequisite:  3113 or permission of instructor.  The course focus is on the molecular and cellular bases of disease and alterations in cellular process that lead to the development of various pathological conditions.  Topics include symptoms of cellular disease, pathology of organelles, cell injury, cell death, immunopathology, neoplasia and genetic disorders.  No student may earn credit for both 4113 and 5113. (F)

5153 Endocrinology (Slashlisted with 4153). Prerequisite: 3103; 3113 strongly recommended. The major emphasis of the course is how hormones work at the molecular, cellular, and physiological levels. The course examines the molecular biology of hormonal mechanisms and the integration of those mechanisms to maintain crucial homeostatic control systems. Hormones and their receptors are considered in terms of chemistry, biosynthesis, and mechanisms of action. The course focuses primarily on vertebrate hormones. No student may earn credit for both 4153 and 5153. (Sp)

5172 Cellular and Molecular Techniques (Slashlilsted with 4172). Prerequisite: ZOO 3113 or graduate standing or permission of instructor. A hands-on introduction to molecular techniques used in modern cell/molecular laboratories and discussed in cell biology courses. No student may earn credit for both 4172 and 5172. (Irreg.)

5193 Life History (Slashlisted with 4193). Prerequisite: graduate standing or permission of instructor. Overview of theory and empirical studies of life history evolution. Includes lecture, discussion of the primary literature, and analysis of some basic life history problems. No student may earn credit for both ZOO 4193 and 5193 (Irreg.)

5204 Vertebrate Paleobiology (Crosslisted with Geology 5204; Slashlisted with 4204). Prerequisite: graduate standing and permission of instructor. Systematics, relationships, zoogeography and evolutionary morphology of the major groups of vertebrates. Field trips. No student may earn credit for both 4204 and 5204. (Irreg.)

5223 Cellular and Molecular Neurobiology (Slashlisted with 4223). Prerequisites: ZOO 3103, ZOO 3113, ZOO 3203, or ZOO 3333, or permission of instructor. Introduction to cellular and molecular neurobiology through reading and discussion of landmark research papers in five current areas of research. Emphasis will be on understanding modern cellular and molecular research methods and on critical interpretation of scientific data. No student may earn credit for both 4223 and 5223. (Irreg.)

5233 Neurobiology of Disease (Slashlisted with 4233). Prerequisite: ZOO 4833 (preferred), or ZOO 3113 and ZOO 3333, or ZOO 2124 or ZOO 3103; or permission of the instructor. Cellular and molecular mechanisms underlying both normal neuronal function and neuronal disorders. Includes a review of basic concepts in neuroscience through traditional lectures, and reading and discussion of original research articles. Students are required to give oral presentations, write critiques and term papers, and present research posters. No student may earn credit for both ZOO 4233 and ZOO 5233. (F)

5253 Evolution of Development (Slashlisted with 4253). Prerequisite: permission of the instructor. Introduction to evolutionary developmental biology (EVO DEVO), study of the developmental basis of evolutionary changes and how developmental mechanisms and developmentally important genes evolve. Latest techniques and methods in evolutionary biology, embryology, developmental biology, molecular genetics, population biology, phylogenetics and genomics. No student may earn credit for both 4253 and 5253. (Irreg.)

5293 Cytology Ultrastructure (Crosslisted with Botany, Microbiology 5293). Prerequisite: 12 hours of biology. A descriptive survey of bacterial, plant and animal cells. Emphasis will be given to the ultrastructural morphology of cellular organelles and their functional significance. (Irreg.)

5333 Ecological Genetics. Prerequisite: 3333 required; 3013 recommended. History of population and ecological genetics; genetic systems; emphasis on experimental aspects of population genetics and measurement of selection and levels of genetic variation in different ecological situations; development of quantitative characters; genetic load; drift; enzyme variation; chromosome variation. (Irreg.)

5343 Developmental Genetics. Prerequisite: 3333. Covers the regulatory control of development in simple viral and bacterial operons, but emphasizes eukaryotic development and genetic organization. Topics will include DNA and chromosome structure, intron processing, nuclear-cytoplasmic interaction, pattern formation and aging. (Irreg.)

5353 Molecular Techniques for Field Biology (Slashlisted with 4353). Prerequisite: permission of instructor; 3333 or 3403 recommended. Selected protocols and data interpretation using molecular techniques to study protein and DNA variation in natural populations and the application of molecular techniques to research problems in ecology, systematics, animal behavior, conservation biology, and related areas. Graduate students enrolled in 5353 will have additional project expectations and written work. Taught at the OU Biological Station. No student may earn credit for both 4353 and 5353. Laboratory (Su)

5364 Transmission Electron Microscopy (Crosslisted with Botany 5364, Microbiology 5364). Prerequisite: permission of instructor. Introduction to the theory of transmission electron microscopy and practical instruction in specimen preparation, ultramicrotomy, instrument operation, photography and quantitative methods. Laboratory. (F)

5374 Scanning Electron Microscopy (Crosslisted with Botany 5374, Microbiology 5374). Prerequisite: basic chemistry; basic physics; demonstrated need; permission of instructor. Principles of scanning electron microscopy are combined with training in the operation of the SEM and ancillary equipment. Students will be certified in the operation of all equipment. Sample preparationon a variety of samples and darkroom proedures will be performed. Independent project with oral report and poster required. Laboratory (Irreg.)

5383 Functional Genomics (Slashlisted with 4383). Prerequisite: permission of the instructor. Introduction to how to study the functions (broadly including biochemical activity, cell biological function or organismal function) of all the genes in a genome. Latest techniques and methods in genetics, molecular biology, microarrays, proteomics and computation biology. No student may earn credit for both 4383 and 5383.(Irreg.)

5403 Population Ecology. Prerequisite: graduate standing. History, demography, environmental factors, density-dependent factors, genetics and population ecology, theories of population and community organization (ideas of Elton, Williams, Preston, MacArthur, Smith, Hairston, and Slodbodkin). No laboratory. (Irreg.)

5413 Community Ecology. Prerequisite: 3403 and Mathematics 1743 or 1823, or permission. Theoretical and empirical study of the structure and organization of natural communities. Topics include competition, predation, disturbance, abiotic gradients, species equilibria.

5423 Stream Ecology (Slashlisted with 4423). Prerequisite: graduate standing or permission of instructor. A combined lecture/laboratory course that focuses on the physical, chemical, and biological features of stream ecosystems, including current theories explaining species interactions and stream function. Course requirements/evaluation including a midterm and final examination, individual research papers and presentations, participation in group laboratory and field experiments, and reading and discussing the primary literature. No student may earn credit for both 4423 and 5423. Laboratory (F-odd numbered years)

5443 Physiological Ecology. Prerequisite: permission of instructor. A study of the physiological adjustments made by animals to changes in their external environment. (Irreg.)

5453 Advanced Ecology and Evolutionary Biology (Crosslisted with Botany and Microbiology). Prerequisite: ZOO 3403. Required for students in the ecology and evolutionary biology doctoral program. An introduction to current research opportunities and research programs in ecology and evolutionary biology at the University of Oklahoma. Specific topics and lecturers will vary from week to week to give students a broad overview of ongoing research projects. (F)

5464 Molecular Methods in Ecology and Evolution. Prerequisite: Permission of the instructor. Laboratory-based course designed to develop skills commonly used in molecular population genetics, ecology and systematics. Emphasis will be on DNA-based methods such as PCR, DNA sequencing, and microsatellite DNA procedures. Theoretical, physical and biochemical bases of methods will be stressed. Appropriate applications of methods and basic data analysis will also be covered. Laboratory (Irreg.)

5471 Seminar in Ecology and Evolutionary Biology (Crosslisted with Botany and Microbiology). Prerequisite: graduate standing. Two semesters of enrollment are required for students in the ecology and evolutionary biology doctoral program. An intensive, student-based seminar in which students present both proposals and ongoing progress reports on doctoral level research projects in ecology and evolutionary biology. (F, Sp)

5483 Fish Ecology. Prerequisite: ZOO 1124, or ZOO 1114 and ZOO 1121; and BIOL 1134, or permission of instructor. Emphasizes fundamental ecology of freshwater and marine fishes worldwide. Focus will be on the historical literature and development of fish ecology, as well as modern comparative and experimental approaches in distributional ecology, fish community structure/function, abiotic and biotic interactions affecting fishes, and direct and indirect effects of fishes in ecosystems. (Irreg.)

5494 Biology of Fishes. Prerequisite: ZOO 4463 or permissio of instructor. Natural history and ecology of freshwater fishes. Field trips. Laboratory. (F odd-numbered years)

5503 An Introduction to Fish Culture. Prerequisite: ZOO 4463 or permissio of instructor. Principles of warm water fish culture, practical culture and field trips. Laboratory. (Irreg.)

5563 Biological Conservation (Slashlisted with 4563). Prerequisite: permission of instructor. The human economy currently entrains more than 40% of global net primary production. This human demand for energy and the effluent subsequently produced have pervasive effects on natural systems and human welfare. This course uses primary literature to guide discussion of ecological, legal, and societal issues affecting biodiversity in North America and globally. Topics range from organism centered approaches to conservation to broader implications of alterations of global processes such as elemental cycles and atmospheric temperatures. No student may earn credit for both 4563 and 5563. (F-even numbered years)

5573 Conservation Genetics (Slashlisted with 4573). Prerequisite: 3333 or permission of the instructor. This lecture/discussion course will examine the use of population genetic/ecological genetic principles in the study and management of populations of threatened and/or endangered species. No student may earn credit for both 4573 and 5573. (Sp)

5663 Advanced Limnology (Slashlisted with 4663). Prerequisite: 4423 or 5423, or 4463 and 4471, or permission of instructor. May be repeated with change of content; maximum credit 6 hrs. detailed study of fundamental or contemporary topics within limnology, such as Biogeochemistry, nutrient cycling, ecological stoichiometry, biodiversity, and predator-prey and food-web dynamics in aquatic communities. No student may earn credit for both 4663 and 5663 on the same topic. (Sp)

5693 Fishery Management. Prerequisite: 4462, 5494 or permission of instructor. Relationship of ecological principals to the management of fisheries. Influence of physico-chemical factors in the environment will be emphasized. A combination of field and laboratory exercises will relate basic fishery biology techniques to applications in management. (Irreg.)

5753 Principles of Phylogenetics (Slashlisted with 4753). Prerequisite: ZOO 2013 and ZOO 3333. Theory and practice of inferring evolutionary history from molecular and morphological data. Applications of the phylogenetic approach in systematics, comparative biology, molecular evolution, and genomics will be covered. No student may earn credit for both 4753 and 5753. (Irreg.)

5813 Hormones and Behavior (Slashlisted with 4813). Prerequisite: 3083 or 3103 or permission of instructor. The relationship between hormones and behavior from an evolutionary perspective. Focus on reproductive and social behavior, with coverage of biological rhythms, food and water intake, and learning. No student may earn credit for both 4813 and 5813. (Irreg.)

5833 Introduction to Neurobiology (Slashlisted with 4833). Prerequisite: permission of instructor. Introduction to cellular and behavioral neurobiology. Topics covered will include cellular neurobiology, neurophysiology, neuroanatomy, sensory processing, movement, and neurobiology of behavior. No student may earn credit for both 4833 and 5833. (F)

5843 Introduction to Molecular Biology (Crosslisted with Botany and Microbiology; Slashlisted with 4843). Prerequisite: permission of instructor. Introduction to the characteristics and biological functions of nucleic acids and proteins in living cells with emphasis on nucleic acid replication, transcription, translation and regulation; also emphasis on the molecular aspects of microbial genetics—transformation, transduction and conjugation; and emphasis on molecular immunology and genetic engineering/recombinant DNA technology. No student may earn credit for both 4843 and 5843. (F)

5863 Neural Control of Movement (Slashlisted with 4863). Prerequisite: ZOO 3103 or ZOO 3113 or ZOO 4833/5833 or permission of instructor. Introduction to neural control of movement through reading and discussion of key original research articles from the 19th century to the present. Students lead discussions and write essays addressing a general question, utilizing data from the articles; students in 5863 write an additional essay. Topics include localization of function, sensory vs. central contributions, roles of single neurons, effects of neuromodulators, and motor learning. No student may earn credit for both 4863 and 5863. (Sp)

5871 Current Topics in Neurobiology (Slashlisted with 4871).  Prerequisite:  Permission of the instructor.  May be repeated; maximum credit three hours.  A seminar course designed to develop a student’s abilities to interpret and critically evaluate research in cellular and behavioral neurobiology.  Involves both public seminars and journal club style discussions of contemporary literature.  No student may earn credit for both 4871 and 5871 concurrently.  (F, Sp)

5893 Behavioral Neurobiology (Slashlisted with 4893). Prerequisite: ZOO 3103 or ZOO 3113, or ZOO 4833/ZOO 5833, or permission of instructor. Examines neurobiological mechanisms of natural animal behaviors (i.e. neuroethology), utilizing textbook and lectures as well as in-depth reading, discussion, and student presentation of original research articles. No student may earn credit for both 4893 and 5893. (Irreg.)

5903 Bioinformatics: Applications. Prerequisites: Botany/Zoology 3333 or 3113, or Botany/Microbiology/Zoology 4843/5843, or Microbiology 3813, or Chemistry 3653 or 5853, or equivalent and basic statistics (Psychology 2003 or 2113, or Zoology 2913, or Biostatistics and Epidemiology 5163, or equivalent) and permission of instructor. This course is designed to provide an understanding of theory and application of computational methods for molecular biology research. The course will cover the following topics: Linux basics, genome sequencing and databases, sequence alignment, database searching for similar sequences, gene analysis, protein analysis, genome analysis, phylogenetic prediction, and a brief introduction to PERL programming. (Irreg.)

5913 Bioinformatics: Programming. Prerequisites: 5903 and permission of instructor. This course is designed to provide programming skills needed to perform large-scale genomic analysis in research and professional settings. The Perl programming language will be taught by introducing features of the Perl programming language in the context of and with applications to bioinformatics. (Irreg.)

5970 Special Topics in Zoology. 1 to 3 hours. Prerequisite: permission of instructor. May be repeated with change of topic; maximum credit 12 hours. Special topics course; may include laboratory or field work. (Sp, Su)

5980 Research for Master's Thesis. Variable enrollment, two to nine hours; maximum credit applicable toward degree, six hours. Laboratory (F, Sp, Su)

5990 Independent Study. 1 to 6 hours. Prerequisite: permission of instructor. May be repeated in different fields; maximum credit 12 hours. Directed readings. Written report required. No laboratory. (F, Sp, Su)

6003 Ecological Modeling (Crosslisted with Botany and Microbiology 6003). Prerequisite: one computer course, one course in ecology, or permission of instructor. Trains students to use modeling tools in their research and to gain greater ability to understand, appreciate, and criticize modeling work. Students will learn general procedure and principles with case studies of successful models in ecology and participate in course projects to gain hands-on experience in model development. (Irreg.)

6083 Evolution of Social Behavior. Prerequisite: 3083 and permission of instructor. History of sociobiological concepts; relationships of ecology, ethology, and evolution. (Irreg.)

6152 Methods in Endocrinology. Prerequisite: 5153 or concurrent enrollment in 5153, permission. Experiments are designed to illustrate the modern as well as classical techniques and instrumentations used in endocrinology. Students are responsible for post-operative care and treatment of animals. An individual problem is chosen by each student. Laboratory (Irreg.)

6213 Developmental Behavioral Neuroscience. Prerequisite: graduate standing or permission of instructor. Discussion of specific developmental issues from both the behavioral and neurobiological perspectives of analysis. Can not be taken for credit if 4213 has been previously taken. (Irreg.)

6970 Seminar. 1 to 4 hours. Prerequisite: graduate standing and permission of instructor. May be repeated; maximum credit 12 hours; no more than six hours may be in any one field. No laboratory. (F, Sp, Su)

6980 Research for Doctoral Dissertation. (F, Sp, Su)

Updated: February 20, 2020